Dr Simon Loader

Grants

Molecular tools for uncovering Vertebrate diversity knowledge: The history of the enigmatic “Bogota” bird skins’
Role: Principal investigatorLoader SP
Funding: Department for Business, Innovation & Skills
Total value £41,400 (to Museum £41,400)
Dates: 2018 - 2018

Hidden affiliate diversity and threats
Role: Principal investigatorOkamura B
Co-investigator: Bass D
Funding: Leverhulme Trust
Dates: 2020 - 2023

CHROMREP - An integrative approach linking chromosomal evolution and biodiversity in reptiles from Madagascar
Role: Principal investigatorMezzasalma M
Co-investigator: Streicher J
Dates: 2018 - 2020

Fellowships

UKPSF and Fellowship of the Higher Education Authority,

Honorary Fellow, London, United Kingdom,

Committees

Chair, Amphibian Specialist Group Regional Chair for East Africa, IUCN, 2008 - on going.

Memberships

Council member, 2013 - 2016.

Editorships

Associate editor, Bulletin- British Herpetological Society, 2011 - on going.

Publications

Liedtke HC, Lyakurwa JV, Lawson LP, Menegon M, Garrido-Priego M, Mariaux J, Ngalason W, Channing A, Owen NR, Bittencourt-Silva GB, Wilkinson M, Larson JG, Gvoždík V, Loader SP (2022) Thirty years of amphibian surveys in the Ukagurus Mountains of Tanzania reveal new species, yet others are in decline. African Journal of Herpetology, 71 (2) : 119 - 138. doi: 10.1080/21564574.2022.2043945

Liedtke HC, Soler-Navarro DJ, Gomez-Mestre I, Loader SP, Rödel M-O (2021) Parallel diversification of the African tree toad genus Nectophryne (Bufonidae). Molecular Phylogenetics and Evolution, 162 : 107184 - 107184. doi: 10.1016/j.ympev.2021.107184

Ngwava JM, Barratt CD, Boakes E, Bwong BA, Channing A, Couchman O, Lötters S, Malonza PK, Muchai V, Nguku JK, Nyamache J, Owen N, Wasonga V, Loader SP (2021) Species-specific or assemblage-wide decline? The case of Arthroleptides dutoiti Loveridge, 1935 and the amphibian assemblage of Mount Elgon, Kenya. African Journal of Herpetology, 70 (1) : 53 - 60. doi: 10.1080/21564574.2021.1891977

Bond AL, Taylor C, Kinchin-Smith D, Fox D, Witcutt E, Ryan PG, Loader SP, Weimerskirch H (2021) A juvenile Tristan albatross (Diomedea dabbenena) on land at the Crozet Islands. Polar Biology, 44 (1) : 229 - 233. doi: 10.1007/s00300-020-02786-0

Mezzasalma M, Guarin FM, Loader SP, Odierna G, Streicher JW, Cooper N (2020) First karyological analysis of the endemic Malagasy phantom gecko Matoatoa brevipes (Squamata: Gekkonidae). ACTA HERPETOLOGICA, 15 (2) : 137 - 141. doi: 10.13128/a_h-8437

Bwong BA, Barratt CD, Loader SP, Nyamache JO, Menegon M, Malonza PK, Nagel P, Lawson LP (2020) Systematics of Hyperolius mitchelli Loveridge, 1953 and Hyperolius rubrovermiculatus Schiøtz, 1975 (Anura: Hyperoliidae) and assignment of populations from northern Tanzania. African Journal of Herpetology, 69 (2) : 165 - 181. doi: 10.1080/21564574.2020.1768155

Greenwood L, Loader SP, Lawson L, Greenbaum E, Zimkus BM (2020) A new species of Phrynobatrachus (Amphibia: Anura: Phrynobatrachidae) from the Northern Mountains of Tanzania. Journal of Natural History, 54 (1-4) : 63 - 85. doi: 10.1080/00222933.2020.1757171

Streicher JW, Loader SP, Varela-Jaramillo A, Montoya P, de Sá RO (2020) Analysis of ultraconserved elements supports African origins of narrow-mouthed frogs. Molecular Phylogenetics and Evolution, 146 : 106771 - 106771. doi: 10.1016/j.ympev.2020.106771

Streicher JW, Sadler R, Loader SP (2020) Amphibian taxonomy: early 21st century case studies. Journal of Natural History, 54 (1-4) : 1 - 13. doi: 10.1080/00222933.2020.1777339

BRANCH WR, BAYLISS J, BITTENCOURT-SILVA GB, CONRADIE W, ENGELBRECHT HM, LOADER SP, MENEGON M, NANVONAMUQUITXO C, TOLLEY KA (null) A new species of tree snake (Dipsadoboa, Serpentes: Colubridae) from ‘sky island’ forests in northern Mozambique, with notes on other members of the Dipsadoboa werneri group. Zootaxa, 4646 (3) : 541 - 563. doi: 10.11646/zootaxa.4646.3.6

Portik DM, Bell RC, Blackburn DC, Bauer AM, Barratt CD, Branch WR, Burger M, Channing A, Colston TJ, Conradie W, Dehling JM, Drewes RC, Ernst R, Greenbaum E, Gvoždík V, Harvey J, Hillers A, Hirschfeld M, Jongsma GFM, Kielgast J, Kouete MT, Lawson LP, Leaché AD, Loader SP, Lötters S, Meijden AVD, Menegon M, Müller S, Nagy ZT, Ofori-Boateng C, Ohler A, Papenfuss TJ, Rößler D, Sinsch U, Rödel M-O, Veith M, Vindum J, Zassi-Boulou A-G, McGuire JA (2019) Sexual Dichromatism Drives Diversification within a Major Radiation of African Amphibians. Systematic Biology, 68 (6) : 859 - 875. doi: 10.1093/sysbio/syz023

Crotti M, Barratt CD, Loader SP, Gower DJ, Streicher JW (2019) Causes and analytical impacts of missing data in RADseq phylogenetics: Insights from an African frog (Afrixalus ). Zoologica Scripta, 48 (2) : 157 - 167. doi: 10.1111/zsc.12335

Barratt CD, Bwong BA, Jehle R, Liedtke HC, Nagel P, Onstein RE, Portik DM, Streicher JW, Loader SP (2018) Vanishing refuge? Testing the forest refuge hypothesis in coastal East Africa using genome-wide sequence data for seven amphibians. Molecular Ecology, 27 (21) : 4289 - 4308. doi: 10.1111/mec.14862

Conradie W, Bittencourt-Silva GB, Farooq HM, Loader SP, Menegon M, Tolley KA (2018) New species of Mongrel Frogs (Pyxicephalidae: Nothophryne) for northern Mozambique inselbergs. African Journal of Herpetology, 67 (1) : 61 - 85. doi: 10.1080/21564574.2017.1376714

Bittencourt-Silva GB, Lawson LP, Tolley KA, Portik DM, Barratt CD, Nagel P, Loader SP (2017) Impact of species delimitation and sampling on niche models and phylogeographical inference: A case study of the East African reed frog Hyperolius substriatus Ahl, 1931. Molecular Phylogenetics and Evolution, 114 : 261 - 270. doi: 10.1016/j.ympev.2017.06.022

WILKINSON M, MALONZA PK, CAMPBELL P, LOADER SP (null) A new species of Boulengerula Tornier, 1896 (Amphibia: Gymnophiona: Herpelidae) from Kenya and the “rediscovery” of Boulengerula denhardti. Zootaxa, 4286 (4) : 525 - 525. doi: 10.11646/zootaxa.4286.4.5

Liedtke HC, Müller H, Hafner J, Penner J, Gower DJ, Mazuch T, Rödel M-O, Loader SP (2017) Terrestrial reproduction as an adaptation to steep terrain in African toads. Proceedings of the Royal Society B: Biological Sciences, 284 (1851) : 20162598 - 20162598. doi: 10.1098/rspb.2016.2598

Barratt CD, Lawson LP, Bittencourt-Silva GB, Doggart N, Morgan-Brown T, Nagel P, Loader SP (2017) A new, narrowly distributed, and critically endangered species of spiny-throated reed frog (Anura: Hyperoliidae) from a highly threatened coastal forest reserve in Tanzania. HERPETOLOGICAL JOURNAL, 27 (1) : 13 - 24.

Liedtke HC, Müller H, Rödel M-O, Menegon M, Gonwouo LN, Barej MF, Gvoždík V, Schmitz A, Channing A, Nagel P, Loader SP (2016) No ecological opportunity signal on a continental scale? Diversification and life-history evolution of African true toads (Anura: Bufonidae). Evolution, 70 (8) : 1717 - 1733. doi: 10.1111/evo.12985

Geiser MF, Hagmann R, Nagel P, Loader SP (2016) A first broad-scale molecular phylogeny of Prionoceridae (Coleoptera: Cleroidea) provides insight into taxonomy, biogeography and life history evolution. ARTHROPOD SYSTEMATICS & PHYLOGENY, 74 (1) : 3 - 21.

Bittencourt-Silva GB, Conradie W, Siu-Ting K, Tolley KA, Channing A, Cunningham M, Farooq HM, Menegon M, Loader SP (2016) The phylogenetic position and diversity of the enigmatic mongrel frog Nothophryne Poynton, 1963 (Amphibia, Anura). Molecular Phylogenetics and Evolution, 99 : 89 - 102. doi: 10.1016/j.ympev.2016.03.021

Beck J, Liedtke HC, Widler S, Altermatt F, Loader SP, Hagmann R, Lang S, Fiedler K (2016) Patterns or mechanisms? Bergmann’s and Rapoport’s rule in moths along an elevational gradient. Community Ecology, 17 (2) : 137 - 148. doi: 10.1556/168.2016.17.2.2

Conradie W, Bittencourt-Silva G, Engelbrecht HM, Loader SP, Menegon M, Nanvonamuquitxo C, Scott M, Tolley KA (null) Exploration into the hidden world of Mozambique’s sky island forests: new discoveries of reptiles and amphibians. Zoosystematics and Evolution, 92 (2) : 163 - 180. doi: 10.3897/zse.92.9948

CONRADIE W, BITTENCOURT-SILVA GB, LOADER SP, MENEGON M, NANVONAMUQUITXO C, KOTZÉ A, DALTON DL, ENGELBRECHT HM, TOLLEY KA (2016) Batrachochytrium dendrobatidis Survey of Amphibians in the Northern Mozambique “Sky Islands” and Low-lying Areas. Herpetological Review, 47 (1) : 42 - 46.

Gereau RE, Cumberlidge N, Hemp C, Hochkirch A, Jones T, Kariuki M, Lange CN, Loader SP, Malonza PK, Menegon M (2016) Globally threatened biodiversity of the Eastern Arc mountains and coastal forests of Kenya and Tanzania. Journal of East African Natural History, 105 (1) : 115 - 201.

Hertach T, Puissant S, Gogala M, Trilar T, Hagmann R, Baur H, Kunz G, Wade EJ, Loader SP, Simon C, Nagel P (null) Complex within a Complex: Integrative Taxonomy Reveals Hidden Diversity in Cicadetta brevipennis (Hemiptera: Cicadidae) and Unexpected Relationships with a Song Divergent Relative. PLOS ONE, 11 (11) : e0165562 - e0165562. doi: 10.1371/journal.pone.0165562

Poynton JC, Loader SP, Conradie W, Rödel M-O, Liedtke HC (2016) Designation and description of a neotype of Sclerophrys maculata (Hallowell, 1854), and reinstatement of S. pusilla (Mertens, 1937) (Amphibia: Anura: Bufonidae).. Zootaxa, 4098 (1) : 73 - 94. doi: 10.11646/zootaxa.4098.1.3

Lawson LP, Bates JM, Menegon M, Loader SP (2015) Divergence at the edges: peripatric isolation in the montane spiny throated reed frog complex. BMC Evolutionary Biology, 15 (1) : doi: 10.1186/s12862-015-0384-3

Loader SP, Lawson LP, Portik DM, Menegon M (2015) Three new species of spiny throated reed frogs (Anura: Hyperoliidae) from evergreen forests of Tanzania. BMC Research Notes, 8 (1) : doi: 10.1186/s13104-015-1050-y

Loader SP (2015) The distribution of Mertensophryne cf. anotis with a new record in Northern Mozambique. Herpetology notes, 8 : 305 - 307.

Loader SP (2015) Stormy waters: Review of The Monkey's Voyage, by Alan de Queiroz, Basic Books, pp. 368. :

Menegon M, Loader SP, Davenport TRB, Howell KM, Tilbury CR, Machaga S, Tolley KA (2015) A new species of Chameleon (Sauria: Chamaeleonidae: Kinyongia) highlights the biological affinities between the Southern Highlands and Eastern Arc Mountains of Tanzania. ACTA HERPETOLOGICA, 10 (2) : 111 - 120.

Penske S, Gvozdik V, Menegon M, Loader SP, Muller H (2015) Description of the tadpole of Leptopelis cf. grandiceps (Amphibia: Anura: Arthroleptidae) from the Uluguru Mountains, Tanzania. HERPETOLOGICAL JOURNAL, 25 (1) : 61 - 64.

Menegon M, Loader SP, Marsden SJ, Branch WR, Davenport TRB, Ursenbacher S (2014) The genus Atheris (Serpentes: Viperidae) in East Africa: Phylogeny and the role of rifting and climate in shaping the current pattern of species diversity. Molecular Phylogenetics and Evolution, 79 : 12 - 22. doi: 10.1016/j.ympev.2014.06.007

Loader SP, Sara Ceccarelli F, Menegon M, Howell KM, Kassahun R, Mengistu AA, Saber SA, Gebresenbet F, de Sá R, Davenport TRB, Larson JG, Müller H, Wilkinson M, Gower DJ (2014) Persistence and stability of Eastern Afromontane forests: evidence from brevicipitid frogs. Journal of Biogeography, 41 (9) : 1781 - 1792. doi: 10.1111/jbi.12331

Gvoždík V, Tillack F, Menegon M, Loader SP (2014) The status of Leptopelis barbouri Ahl, 1929 and eleven other nomina of the current tree-frog genus Leptopelis (Arthroleptidae) described from East Africa, with a redescription of Leptopelis grandiceps Ahl, 1929.. Zootaxa, 3793 : 165 - 187. doi: 10.11646/zootaxa.3793.1.8

Ceccarelli FS, Menegon M, Tolley KA, Tilbury CR, Gower DJ, Laserna MH, Kasahun R, Rodriguez-Prieto A, Hagmann R, Loader SP (2014) Evolutionary relationships, species delimitation and biogeography of Eastern Afromontane horned chameleons (Chamaeleonidae: Trioceros). Molecular Phylogenetics and Evolution, 80 : 125 - 136. doi: 10.1016/j.ympev.2014.07.023

Liedtke HC, Hügli D, Dehling JM, Pupin F, Menegon M, Plumptre AJ, Kujirakwinja D, Loader SP (2014) One or two species? On the case of Hyperolius discodactylus Ahl, 1931 and H. alticola Ahl, 1931 (Anura: Hyperoliidae).. Zootaxa, 3768 : 253 - 290. doi: 10.11646/zootaxa.3768.3.2

Barej MF, Schmitz A, Günther R, Loader SP, Mahlow K, Rödel M-O (2014) The first endemic West African vertebrate family – a new anuran family highlighting the uniqueness of the Upper Guinean biodiversity hotspot. Frontiers in Zoology, 11 (1) : 8 - 8. doi: 10.1186/1742-9994-11-8

Barej MF, Rödel M-O, Loader SP, Menegon M, Gonwouo NL, Penner J, Gvoždík V, Günther R, Bell RC, Nagel P, Schmitz A (2014) Light shines through the spindrift – Phylogeny of African torrent frogs (Amphibia, Anura, Petropedetidae). Molecular Phylogenetics and Evolution, 71 : 261 - 273. doi: 10.1016/j.ympev.2013.11.001

San Mauro D, Gower DJ, Müller H, Loader SP, Zardoya R, Nussbaum RA, Wilkinson M (2014) Life-history evolution and mitogenomic phylogeny of caecilian amphibians. Molecular Phylogenetics and Evolution, 73 : 177 - 189. doi: 10.1016/j.ympev.2014.01.009

Barratt CD, Tonelli E, Menegon M, Doggart N, Bowkett A, Harris WE, Howell K, Ngalason W, Loader SP (2014) Fragmented habitats and species: the challenges of Amphibian conservation in Tanzania today. Froglog,

Bwong BA, Malonza PK, Wasonga DV, Nyamache JO, Nagel P, Loader SP (2014) At a Biogeographical Crossroads: Amphibian Paradise in Shimba Hills of Kenya. Froglog, 72 - 74.

Ceccarelli FS, Menegon M, Tolley KA, Tilbury CR, Gower DJ, Laserna MH, Kasahun R, Rodriguez-Prieto A, Hagmann R, Loader SP (2014) Evolutionary relationships, species delimitation and biogeography. null,

Liedtke HC, Müller H, Hafner J, Nagel P, Loader SP (2014) Interspecific patterns for egg and clutch sizes of African Bufonidae (Amphibia: Anura). Zoologischer Anzeiger - A Journal of Comparative Zoology, 253 (4) : 309 - 315. doi: 10.1016/j.jcz.2014.02.003

Mapouyat L, Hirschfeld M, Rödel M, Liedtke HC, Loader SP, Gonwouo LN, Dahmen M, Doherty-Bone TM, Barej MF (2014) The tadpoles of nine Cameroonian Leptodactylodon species (Amphibia, Anura, Arthroleptidae). Zootaxa, 3765 (1) : 029 - 053.

Mapouyat L, Hirschfeld M, Rödel M-O, Liedtke HC, Loader SP, Gonwouo LN, Dahmen M, Doherty-Bone TM, Barej MF (null) <p class="HeadingRunIn"><strong>The tadpoles of nine Cameroonian <em>Leptodactylodon</em> species (Amphibia, Anura, Arthroleptidae)</strong></p>. Zootaxa, 3765 (1) : 29 - 29. doi: 10.11646/zootaxa.3765.1.2

Siu-Ting K, Gower DJ, Pisani D, Kassahun R, Gebresenbet F, Menegon M, Mengistu AA, Saber SA, de Sá R, Wilkinson M, Loader SP (2014) Evolutionary relationships of the Critically Endangered frog Ericabatrachus baleensisLargen, 1991 with notes on incorporating previously unsampled taxa into large-scale phylogenetic analyses. BMC Evolutionary Biology, 14 (1) : doi: 10.1186/1471-2148-14-44

Müller H, Liedtke HC, Menegon M, Beck J, Ballesteros-Mejia L, Nagel P, Loader SP (2013) Forests as promoters of terrestrial life-history strategies in East African amphibians. Biology Letters, 9 (3) : 20121146 - 20121146. doi: 10.1098/rsbl.2012.1146

Gower DJ, Doherty-Bone T, Loader SP, Wilkinson M, Kouete MT, Tapley B, Orton F, Daniel OZ, Wynne F, Flach E, Müller H, Menegon M, Stephen I, Browne RK, Fisher MC, Cunningham AA, Garner TWJ (2013) Batrachochytrium dendrobatidis Infection and Lethal Chytridiomycosis in Caecilian Amphibians (Gymnophiona). EcoHealth, 10 (2) : 173 - 183. doi: 10.1007/s10393-013-0831-9

Loader SP, Ceccarelli FS, Wilkinson M, Menegon M, Mariaux J, de Sá RO, Howell KM, Gower DJ (2013) Species boundaries and biogeography of East African torrent frogs of the genus Petropedetes (Amphibia: Anura: Petropeditidae). African Journal of Herpetology, 62 (1) : 40 - 48. doi: 10.1080/21564574.2013.781549

Wilkinson M, Gower DJ, Loader SP, Müller H (2013) A visceral synapomorphy of Scolecomorphus Boulenger, 1883 (Amphibia: Gymnophiona: Scolecomorphidae). African Journal of Herpetology, 62 (1) : 21 - 27. doi: 10.1080/21564574.2013.781548

Doherty-Bone TM, Gonwouo NL, Hirschfeld M, Ohst T, Weldon C, Perkins M, Kouete MT, Browne RK, Loader SP, Gower DJ, Wilkinson MW, Rödel MO, Penner J, Barej MF, Schmitz A, Plötner J, Cunningham AA (2013) Batrachochytrium dendrobatidis in amphibians of Cameroon, including first records for caecilians. Diseases of Aquatic Organisms, 102 (3) : 187 - 194. doi: 10.3354/dao02557

Gower DJ, Aberra ROMANK, Schwaller S, Largen MJ, Collen B, Spawls S, Menegon M, Zimkus BM, de Sá R, Mengistu AA (2013) Long-term data for endemic frog genera reveal potential conservation crisis in the Bale Mountains, Ethiopia. Oryx, 47 (01) : 59 - 69.

Gower DJ, Aberra RK, Schwaller S, Largen MJ, Collen B, Spawls S, Menegon M, Zimkus BM, de Sá R, Mengistu AA, Gebresenbet F, Moore RD, Saber SA, Loader SP (2013) Long-term data for endemic frog genera reveal potential conservation crisis in the Bale Mountains, Ethiopia. Oryx, 47 (1) : 59 - 69. doi: 10.1017/s0030605311001426

Loader SP, Menegon M, Howell KM, Müller H (2013) A celebration of the works of John Charles Poynton. African Journal of Herpetology, 62 (1) : 1 - 4. doi: 10.1080/21564574.2013.793215

Mengistu AA, Nagel P, Getahun A, Saber SA, Loader SP (2013) Updated review of amphibian diversity, distribution and conservation in Ethiopia. Ethiop J Biol Sci, 12 (1) : 81 - 116.

Gower DJ, Doherty-Bone TM, Aberra RK, Mengistu A, Schwaller S, Menegon M, de Sa R, Saber SA, Cunningham AA, Loader SP (2012) High prevalence of the amphibian chytrid fungus (Batrachochytrium dendrobatidis) across multiple taxa and localities in the highlands of Ethiopia. HERPETOLOGICAL JOURNAL, 22 (4) : 225 - 233.

Barratt CD, Horsburgh GJ, Dawson DA, Gower DJ, Wilkinson M, Loader SP, Jehle R (2012) Characterisation of nine microsatellite loci in the caecilian amphibian Boulengerula uluguruensis (Gymnophiona), and their cross-species utility in three congeneric species. Conservation Genetics Resources, 4 (2) : 225 - 229. doi: 10.1007/s12686-011-9512-6

de Sá RO, Streicher JW, Sekonyela R, Forlani MC, Loader SP, Greenbaum E, Richards S, Haddad CFB (2012) Molecular phylogeny of microhylid frogs (Anura: Microhylidae) with emphasis on relationships among New World genera.. BMC evolutionary biology, 12 : doi: 10.1186/1471-2148-12-241

HIRSCHFELD M, BAREJ MF, LOADER SP, RÖDEL M-O (null) Description of two Werneria tadpoles from Cameroon (Amphibia: Anura: Bufonidae). Zootaxa, 3172 (1) : 65 - 65. doi: 10.11646/zootaxa.3172.1.5

Lawson L, Loader SP, Zimkus BM, Hanken J (2012) Terrestrialization, Miniaturization and Rates of Diversification in African Puddle Frogs (Anura: Phrynobatrachidae). null,

Zimkus BM, Lawson L, Loader SP, Hanken J (2012) Terrestrialization, miniaturization and rates of diversification in African puddle frogs (Anura: Phrynobatrachidae).. PloS one, 7 (4) : doi: 10.1371/journal.pone.0035118

Zimkus BM, Lawson L, Loader SP, Hanken J, Swenson NG (2012) Terrestrialization. Miniaturization and Rates of Diversification in African Puddle Frogs (Anura,

Menegon M, Gower DJ, Loader SP (2011) A remarkable new species of Callulina (Amphibia: Anura: Brevicipitidae) with massive, boldly coloured limb glands. ZOOTAXA, 15 - 26.

GOWER DJ, PAPADOPOULOU A, DOHERTY-BONE TM, PUPIN F, SAN MAURO D, LOADER SP, WILKINSON M (null) The systematics of Boulengerula fischeri (Amphibia: Gymnophiona: Caeciliidae) based on morphological and molecular data. Zootaxa, 2767 (1) : 14 - 14. doi: 10.11646/zootaxa.2767.1.2

Loader SP, Poynton JC, Lawson LP, Blackburn DC, Menegon M (2011) Herpetofauna of Montane Areas of Tanzania. 3. Amphibian Diversity in the Northwestern Eastern Arc Mountains, with the Description of a New Species of Arthroleptis (Anura: Arthroleptidae). Fieldiana Life and Earth Sciences, 4 : 90 - 102. doi: 10.3158/2158-5520-4.1.90

Loader SP, Wilkinson M, Cotton JA, Measey GJ, Menegon M, Howell KM, Mueller H, Gower DJ (2011) Molecular phylogenetics of Boulengerula (Amphibia: Gymnophiona: Caeciliidae) and implications for taxonomy, biogeography and conservation. HERPETOLOGICAL JOURNAL, 21 (1) : 5 - 16.

Menegon M, Bracebridge C, Owen N, Loader SP (2011) Herpetofauna of Montane Areas of Tanzania. 4. Amphibians and Reptiles of Mahenge Mountains, with Comments on Biogeography, Diversity, and Conservation. Fieldiana Life and Earth Sciences, 4 : 103 - 111. doi: 10.3158/2158-5520-4.1.103

Loader SP, Gower DJ, Mueller H, Menegon M (2010) Two new species of Callulina (Amphibia: Anura: Brevicipitidae) from the Nguru Mountains, Tanzania. ZOOTAXA, 26 - 42.

Loader SP, Gower DJ, Ngalason W, Menegon M (2010) Three new species of Callulina (Amphibia: Anura: Brevicipitidae) highlight local endemism and conservation plight of Africa's Eastern Arc forests. ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, 160 (3) : 496 - 514. doi: 10.1111/j.1096-3642.2010.00652.x

Van Bocxlaer I, Loader SP, Roelants K, Biju SD, Menegon M, Bossuyt F (2010) Gradual Adaptation Toward a Range-Expansion Phenotype Initiated the Global Radiation of Toads. Science, 327 (5966) : 679 - 682. doi: 10.1126/science.1181707

Loader S (2010) Amphibians as indicators of change in Ethiopian highlands. null,

Loader S, Nagel P (2010) Angebot primär im Master-Studium/Mainly Master-Level. und Informationen zum Studium,

Biju SD, Van Bocxlaer I, Giri VB, Loader SP, Bossuyt F (2009) Two new endemic genera and a new species of toad (Anura: Bufonidae) from the Western Ghats of India. BMC Research Notes, 2 (1) : 241 - 241. doi: 10.1186/1756-0500-2-241

Loader SP, Poynton JC, Davenport TRB, Roedel M-O (2009) Re-description of the type series of Nectophrynoides viviparus (Bufonidae), with a taxonomic reassessment. ZOOTAXA, 41 - 50.

Van Bocxlaer I, Biju SD, Loader SP, Bossuyt F (2009) Toad radiation reveals into-India dispersal as a source of endemism in the Western Ghats-Sri Lanka biodiversity hotspot. BMC Evolutionary Biology, 9 (1) : doi: 10.1186/1471-2148-9-131

MÜLLER H, WILKINSON M, LOADER SP, WIRKNER CS, GOWER DJ (null) Morphology and function of the head in foetal and juvenile Scolecomorphus kirkii (Amphibia: Gymnophiona: Scolecomorphidae). Biological Journal of the Linnean Society, 96 (3) : 491 - 504. doi: 10.1111/j.1095-8312.2008.01152.x

Loader SP, Menegon M, Müller H, Gower DJ, Wilkinson M, Howell KM, Orton F (2009) Remarkable amphibian diversity in the South Nguru Mountains, Tanzania. Froglog, 8 - 10.

Loader S, Mengistu A, Schwaller S, Gower D, Nagel P, Getahun A, Saber S, Kassahun R (2009) Are Ethiopian highlands changing? Amphibians as ecosystem indicators. Mountain Forum Bulletin 9 (2) : ICIMOD :

LOADER SP, MEASEY GJ, DE SÁ RO, MALONZA PK (null) A new brevicipitid species (Brevicipitidae: Callulina) from the fragmented forests of the Taita Hills, Kenya. Zootaxa, 2123 (1) : 55 - 68. doi: 10.11646/zootaxa.2123.1.3

Menegon M, Burgess N, Loader SP, Doggart N, Owen N (2009) The South Nguru Mountains: a new Jewel in the Eastern Arc crown. Oryx, 43 : 174 - 176.

Poynton J, Loader SP, Menegon M (2008) A new giant species ofArthroleptis(Amphibia: Anura) from the forests of the Nguru Mountains, Tanzania. African Journal of Herpetology, 57 (2) : 63 - 74. doi: 10.1080/21564574.2008.9635569

Poynton JC, Loader SP (2008) Clinal Variation and Its Taxonomic Consequences in the Common Tanzanian Forest Frog, Arthroleptis affinis. Copeia, 2008 (3) : 517 - 526. doi: 10.1643/ch-07-057

Andreone F, Channing A, Drewes R, Gerlach J, Glaw F, Howell K, Largen M, Loader S, Lötters S, Minter L (2008) . AMPHIBIANS OF THE AFROTROPICAL REALM. null,

Loader S, Channing A, Howell K, Pickersgill M, Raxworthy CJ, Glaw F, Minter L, Drewes R, Gerlach J, Largen M (2008) AMPHIBIANS OF THE AFROTROPICAL REALM, In: null, null :

Loader SP, Pisani D, Cotton JA, Gower DJ, Day JJ, Wilkinson M (2007) Relative time scales reveal multiple origins of parallel disjunct distributions of African caecilian amphibians. Biology Letters, 3 (5) : 505 - 508. doi: 10.1098/rsbl.2007.0266

Müller H, Loader SP, Ngalason W, Howell KM, Gower DJ (2007) Reproduction in Brevicipitid Frogs (Amphibia: Anura: Brevicipitidae)—Evidence from Probreviceps M. Macrodactylus. Copeia, 2007 (3) : 726 - 733. doi: 10.1643/0045-8511(2007)2007[726:ribfaa]2.0.co;2

Poynton JC, Loader SP, Sherratt E, Clarke BT (2007) Amphibian Diversity in East African Biodiversity Hotspots: Altitudinal and latitudinal Patterns. Biodiversity and Conservation, 16 (4) : 1103 - 1118. doi: 10.1007/s10531-006-9074-1

Roelants K, Gower DJ, Wilkinson M, Loader SP, Biju SD, Guillaume K, Moriau L, Bossuyt F (2007) Global patterns of diversification in the history of modern amphibians. Proceedings of the National Academy of Sciences, 104 (3) : 887 - 892. doi: 10.1073/pnas.0608378104

Burgess ND, Butynski TM, Cordeiro NJ, Doggart NH, Fjeldså J, Howell KM, Kilahama FB, Loader SP, Lovett JC, Mbilinyi B, Menegon M, Moyer DC, Nashanda E, Perkin A, Rovero F, Stanley WT, Stuart SN (2007) The biological importance of the Eastern Arc Mountains of Tanzania and Kenya. Biological Conservation, 134 (2) : 209 - 231. doi: 10.1016/j.biocon.2006.08.015

Menegon M, Salvidio S, Ngalason W, Loader SP (2007) A new dwarf forest toad (Amphibia : Bufonidae : Nectophrynoides) from the Ukaguru Mountains, Tanzania. ZOOTAXA, 31 - 40.

Poynton JC, Loader SP, Sherratt E, Clarke BT (2007) Amphibian diversity in East African biodiversity hotspots: altitudinal and latitudinal patterns. Vertebrate Conservation and Biodiversity, 277 - 292.

Loader SP, Channing A, Menegon M, Davenport TRB (2006) A new species of Probreviceps (Amphibia : Anura) from the Eastern Arc Mountains, Tanzania. ZOOTAXA, 45 - 60.

Jones DT, Loader SP, Gower DJ (2006) Trophic ecology of East African caecilians (Amphibia: Gymnophiona), and their impact on forest soil invertebrates. Journal of Zoology, 269 (1) : 117 - 126. doi: 10.1111/j.1469-7998.2006.00045.x

Loader SP (2005) Systematics and biogeography of amphibians of the African Eastern Arc mountains. null,

Muller H, Measey GJ, Loader SP, Malonza PK (2005) A new species of Boulengerula tornier (Amphibia : Gymnophiona : Caeciliidae) from an isolated mountain block of the Taita Hills, Kenya. ZOOTAXA, 37 - 50.

Menegon M, Salvidio S, Loader SP (2004) Five new species ofNectophrynoidesNoble 1926 (Amphibia Anura Bufonidae) from the Eastern Arc Mountains, Tanzania. Tropical Zoology, 17 (1) : 97 - 121. doi: 10.1080/03946975.2004.10531201

de Sá RO, Loader SP, Channing A (2004) A New Species of Callulina (Anura: Microhylidae) from the West Usambara Mountains, Tanzania. Journal of Herpetology, 38 (2) : 219 - 224. doi: 10.1670/209-03a

Gower DJ, Rasmussen JB, Loader SP, Wilkinson M (2004) The caecilian amphibian Scolecomorphus kirkii Boulenger as prey of the burrowing asp Atractaspis aterrima Gunther: trophic relationships of fossorial vertebrates. African Journal of Ecology, 42 (2) : 83 - 87. doi: 10.1111/j.1365-2028.2004.00495.x

Loader SP, Poynton JC, Mariaux J (2004) Herpetofauna of Mahenge Mountain, Tanzania: a window on African biogeography. AFRICAN ZOOLOGY, 39 (1) : 71 - 76.

Cohen BL, Wilkinson M, Loader S, Howell K, de Sá RO, Gower D, Clarke BT, Doggart N, Rödel MO (2004) Phylogenetic relationships of African Microhylid frogs inferred from DNA sequences of mitochondrial 12S and 16S ribosomal rRNA genes. Org, Divers Evol 4: 227-235. null,

Gower DJ, Loader SP, Moncrieff CB, Wilkinson M (2004) Niche separation and comparative abundance ofBoulengerula boulengeriandScolecomorphus vittatus(Amphibia: Gymnophiona) in an East Usambara forest, Tanzania. African Journal of Herpetology, 53 (2) : 183 - 190. doi: 10.1080/21564574.2004.9635510

Gower DJ, Loader SP, Wilkinson M (2004) Assessing the conservation status of soil‐dwelling vertebrates: Insights from the rediscovery ofTyphlops uluguruensis(Reptilia: Serpentes: Typhlopidae). Systematics and Biodiversity, 2 (1) : 79 - 82. doi: 10.1017/s1477200004001343

LOADER S (2004) Phylogenetic relationships of African microhylid frogs inferred from DNA sequences of mitochondrial 12S and 16S rRNA genes. Organisms Diversity & Evolution, 4 (4) : 227 - 235. doi: 10.1016/j.ode.2004.01.005

Wilkinson M, Loader SP, Müller H, Gower DJ (2004) Taxonomic status and phylogenetic relationships of Boulengerula denhardtiNieden, 1912 (Amphibia, Gymnophiona, Caeciliidae). Zoosystematics and Evolution, 80 (1) : 41 - 51.

Loader SP, Gower DJ, Wilkinson M (2003) Caecilians: mysterious amphibians of the Eastern Arc Mountains. Arc Journal. 15 : : 3 - 4.

Loader SP, Wilkinson M, Gower DJ, Msuya CA (2003) A remarkable young Scolecomorphus vittatus (Amphibia: Gymnophiona: Scolecomorphidae) from the North Pare Mountains, Tanzania. Journal of Zoology, 259 (1) : 93 - 101. doi: 10.1017/s0952836902003060

Wilkinson M, Loader SP, Gower DJ, Sheps JA, Cohen BL (2003) Phylogenetic relationships of African caecilians (Amphibia: Gymnophiona): Insights from mitochondrial rRNA gene sequences. African Journal of Herpetology, 52 (2) : 83 - 92. doi: 10.1080/21564574.2003.9635483

Gower DJ, Kupfer A, Oommen OV, Himstedt W, Nussbaum RA, Loader SP, Presswell B, Müller H, Krishna SB, Boistel R, Wilkinson M (2002) A molecular phylogeny of ichthyophiid caecilians (Amphibia: Gymnophiona: Ichthyophiidae): out of India or out of South East Asia?. Proceedings of the Royal Society of London. Series B: Biological Sciences, 269 (1500) : 1563 - 1569. doi: 10.1098/rspb.2002.2050

Doggart N, Loader S, Perkin A, Mwangoka M (2002) Ngaramia Riverine Forest A biodiversity survey. Tanzania Forest Conservation Group, 14 (10) :

Wilkinson M, Loader S, Msuya CA, Gower D (2002) A remarkable young Scolecomorphus vittatus (Amphibia: Gymnophiona Scolecomorphidae) from the North Pare Mountains, Tanzania. null,

Krause J, Loader SP, Kirkman E, Ruxton GD (1999) Refuge use by fish as a function of body weight changes. acta ethologica, 2 (1) : 29 - 34. doi: 10.1007/pl00012229

Krause J, Loader SP, McDermott J, Ruxton GD (1998) Refuge use by fish as a function of body length–related metabolic expenditure and predation risks. Proceedings of the Royal Society of London. Series B: Biological Sciences, 265 (1413) : 2373 - 2379. doi: 10.1098/rspb.1998.0586

Barratt CD, Bwong BA, Onstein RE, Rosauer DF, Menegon M, Doggart N, Nagel P, Kissling WD, Loader SP (2017) Environmental correlates of phylogenetic endemism in amphibians and the conservation of refugia in the Coastal Forests of Eastern Africa. Diversity and Distributions, 23 (8) : 875 - 887. doi: 10.1111/ddi.12582

Barratt CD, Lawson LP, Bittencourt-Silva GB, Doggart N, Morgan-Brown T, Nagel P, Loader SP (null) Herpetological Journal FULL PAPER. null,

Krause J, Loader SP (null) J. McDermott & GD Ruxton. 1998. Refuge use by fish as a function of body length-related metabolic expenditure and predation risks. Proc. The Roy. Soc. London Series B 265 : 2373 - 2379.

Mengistu AA, Loader SP (null) Amphibian Conservation in the Ethiopian Highlands: The Case of Endemic Tree Frogs (Leptopelis, Arthroleptidae) and Ridged Frogs (Ptychadena, Ptychadenidae). AMPHIBIAN DIVERSITY, DISTRIBUTION AND CONSERVATION IN THE ETHIOPIAN HIGHLANDS,

Mengistu AA, Nagel P, Getahun A, Saber SA, Loader SP (null) Current Status of Amphibian Diversity, Distribution and Conservation in Ethiopia: with emphasis on Leptopelis and Ptychadena (Anura). AMPHIBIAN DIVERSITY, DISTRIBUTION AND CONSERVATION IN THE ETHIOPIAN HIGHLANDS,

Mengistu AA, Nagel P, Getahun A, Saber SA, Menegon M, Gower D, de Sá R, Loader S (null) Phylogeography of Leptopelis (Arthroleptidae, Amphibia) in the Ethiopian Highlands: Molecular and Morphological Investigation. AMPHIBIAN DIVERSITY, DISTRIBUTION AND CONSERVATION IN THE ETHIOPIAN HIGHLANDS,

Mengistu AA, Nagel P, Getahun A, Saber SA, Zimkus B, Menegon M, Gower D, de Sá R, Loader S (null) Phylogeography of Ptychadena (Ptychadenidae, Amphibia) in the Ethiopian Highlands and Rift Valley: Molecular and Morphological Investigation. AMPHIBIAN DIVERSITY, DISTRIBUTION AND CONSERVATION IN THE ETHIOPIAN HIGHLANDS,